The skulls will award 100 plasteel for the first time you pick them up, and 100 or double a martyr skull's value when a teammate picks it up for the first time (credit: IcemayneXXV, Derek Icelord)

The penance categories divide the missions up by the following zones:
Hab Dreyko - Partner Puzzle
Chasm Station HL-16-11 - Solo Puzzle
Chasm Logistratum - Solo Parkour
Ascension Riser 31 - Partner Puzzle
Vigil Station Oblivium - Solo Parkour
Magistrati Oubliette TM8-707 - Partner Puzzle
Silo Cluster 18-66A - Solo Puzzle
Consignment Yard - Solo Puzzle
Smelter Complex HL17-36 - Solo Parkour
Power Matrix HL-17-36 - Solo Puzzle
Excise Vault Spireside-13 - Partner Puzzle
Refinery Delta - Solo Puzzle
Relay Station TRS-150 - Solo Puzzle
Enclavum Baross - Solo Puzzle
Comms-Plex 154/2f - Solo Puzzle
Archivum Sycorax - Partner Parkour
Mercantile HL-70-04 - Solo parkour
Warren 6-19 - Solo puzzle
Dark Communion - Solo parkour

There will be 5 partner activities and 14 solo activities total.

Finding a skull awards a penance for the map worth 10 points and contributes to the maps exploration completion title

Generally i will try to use green for the "puzzle" path, then white for keys/buttons/skulls, generally intractable type things. I will sometimes try to use blue for shortcuts

Partner Puzzle

Go through the map like normal until you enter this building:

Once inside, go right towards the code or go towards the blue line and jump inside the window.

The code is 213, from here you can either jump through the window on the desk, and into the window with the blue mark, or follow along the green path for the scenic route

Once inside, enter the code 213. This code does not change. The next picture has the buttons labled by the order you press them in: the middle button you press first and it has a 2 on it, the left button you press second and it has a 1 on it, and the right button you press third and it has a 3 on it.

Once the code is entered, one person will hold the lever that iss marked "hold" in the picture. This will power the cells on the left-side of the pictures. A second person will press the button labled next to those cells.

The skull will be in the left cell

Solo Puzzle

Go through the map until you get to this point, go underneath the floor and behind the stairs

Hit the valve, then double back and go towards the set of stairs that goes into the lower room

At the bottom of that staircase, there will be another valve. Turn it and go towards the hallway past another set of stairs

Hit this valve, turn around and go up the stairs we just passed

At the top of the stairs, go to the far wall for the other valve

Hit the valve, then proceed up the stairs again.

Once at the top of the stairs, hit the valve and go into the room on the left of the picture

Once inside, the skull is on the right wall

Solo Parkour

Go through the map until you get to the train tunnel, once you dropdown the first time, take an immediate left, jump up on the small platform on the left and onto the wall section.

Once on the wall section, on the left there will be two small metal platforms, jump from one to the next one, and then jump down next to the lantern on the metal platform.

Once you jump down next to the lantern, you will see another, fallen lantern in the darkness. Jump on the platform that the lantern is on and then jump towards the catwalk that the candles sit on.

Once you are on the catwalk, go forward a little bit and look to the left. Jump towards the lantern on the wooden platform, then walk across the metal beam and jump towards the metal plate. From there you can walk, and follow the platforms towards the candle and the skull.

Partner Puzzle

Proceed past the mid event until you get to this point. Go through the large door.

Once inside, you should see a blue light in a container showing where a power cell is. Grab it and place it in the nearby glowing receptacle.

Once the power cell has been inserted, one person will hold the lever to power the elevator and the other person will activate the elevator to go up.

At the top, you will see a powercell. Throw it down and your partner will place it in the other glowing receptacle.

One person will hold the lever, and the other person will activate the elevator.

Once the elevator reaches the bottom, you can go through it to the skull.

Solo Parkour

Proceed through the level until you get to this bridge, then go right.

Just before the door, there will be a dumpster you can jump on. After jumping on the dumpster, jump onto the container. Walk to the edge of the container, and jump onto the next container

From there you will jump onto a platform with corrugated metal. Jump onto the protruding metal beam, and continue to jump along the protruding metal platforms

Once you get around the corner, youll jump towards a grated metal platform, and then inside the building where the skull is.

Partner Puzzle

Proceed through the mission until you get to this room.

You will press the button on your immediate left. One person will stay here and go to the panel with the five buttons with symbols, and the other person will backtrack to the scan room. The buttons with the symbols can each be pressed once to open a door, press the three buttons that dont have a light next to them to open up the first three doors


Back track to the scan room, next to it is a cell with a hole in the wall that is now accessible, go inside.

The first three doors should be opened if the person pressed each of the indicated three buttons once. The fourth door needs some timing, the button on the right in the picture will light up to indicate which button needs to be pressed in the control room. Try to time it so that the door is open when the control room button is pressed.

Press the button on the right to light up the corresponding button in the control room. Both buttons need to be pressed at roughly the same time

Once you go through the fifth door, the skull is in the room directly ahead. You can take a left and follow the hallway to open up a door leading to the room with the button where your partner will be.

Solo Puzzle

Proceed through the level until this area. You will be moving the platform circled by white along the unbroken tracks. Drop down to the left and go into the side room.

In the side room, there should be a barrel next to a broken railing that leads to some stairs. Go up the barrel, onto the platform, and up the stairs

Hit the buttons in this order: down, right, right, down, left. This order does not change. The skull will be on the platform

Solo Puzzle

Go through the map until you reach the showers. This puzzle will involve turning on the marked showers. The lighting makes the shower symbol not appear if they are off to the side, but they will appear if you look directly at the shower


The first one will be on the immediate left.
The second one will be across from it on the right.


Go forward a bit, turn right and the third shower will be on the right side.


The last skull is to your left when you are looking at the third shower.


The skull will be in the next room to the left

Solo Parkour

Get to this area, and open the door by destroying the corruption

Once the door is open, on your immediate left next to the wall will be a jump up to a catwalk.

Proceed into the room, and then go through the next doorway. Jump across the metal platforms until you get to the room with the mini corruptors.

Once there, shoot the corruptor at the top, walk to the other side of the little room and shoot the two corruptors in the hole

After all three corruptors have been shot, you can walk forward and drop down. The door to the skull will now be accessible.

Solo Puzzle

Go through the map until this point and go inside the siderooms.

Next to the doorway, youll see a barrel and boxes. Climb up them and jump onto the scaffolding

On the scaffolding there will be a dead body with a key in his hand. Grab it and jump down and leave the room.

Go back across the bridge, and enter the side room

Open the locker in the room to reveal the skull. The puzzle here was to follow the blood trail to figure out where the key led to.

Partner Puzzle

Get to this point of the map and continue along the walkway

One player will enter the elevator on the left which will go down a level, and the other player will enter the elevator on the right which will go up to another room.

At the top of the right elevator, go inside the room and grab the power cell.

You will bring the powercell back down to the glowing receptacle which on the floor you started on. It will open a door to reveal the skull

Solo Puzzle

Proceed through the level until this point in the map, near the finale. Enter the room in the corner, through either doorway or from the top.

Open the door in the room, and look above the door on the other side of the small bridge for the symbols.

Input those symbols in the consoles on both sides of the door. Left symbol for the left door, and right symbol for the right door.

Once you have done it, the button next to the door with the symbols above it will light up blue and the skull will be inside.

Solo Puzzle

Get to this point in the map and take a left upon going through the door.

Once you get past the fence doorway, there will be a dumpster on the right. You can push it over to reveal a weight.

This weight will be used at the right side of a door. Go forward in the map a bit, and once you reach the little room ahead of the dumpster with the 3 windows and 2 doorways, take a right and there should be a doorway to your left.

The skull will be past the door which opens once you place the weight on the chain. The puzzle here is that you would see the door first, and the puddle of blood underneath the chain. You would follow the trail of blood to a body beside the dumpster and from there you could find the skull with some investigating.

Solo Puzzle

Get to this point in the map and take a right just after the bridge but before the stairs. In the first picture, the location of the skull is the green arrow on the right side of the picture but we will take the path on the left to get to the puzzle controls.

You will see some valves, theses valves control the pipes on the wall down below. From left to right, spin the valves: 3 times, 1 time, 2 times, 3 times. I have coloured the pipes and put splotches of colour on the valve so you can identify which valve moves which pipe. The order should be the same every time

Once the pipes are lined up, the door below will open up. You can jump over the valves to get down. The skull will be beyond the door.

Solo Puzzle

Go through the map until you get to this room, just before the mid-event. The entrance to the mid-event room is on the right side of the first picture, and at the top where the bot outlines are in the second picture.

Go past the lit up board, and look on the other side of it. In the bottom right corner of the board on the side furthest from where from you dropped in from, there will be two symbols. They have been expertly recreated for visual clarity. Continue moving to the wall opposite of where you dropped in.

There will be a circle of candles and symbols. Find the two corresponding symbols and light them.

Once the candles are lit, a door across from the candles will open. Inside will be the skull

Partner Parkour

Get to this large room, there will be buttons that can be pressed and chandeliers that will raise or lower in response.

We will first lower this chandelier, then someone will get on. The other person will hit the lever and raise the chandelier.

From here you will continue until you get to a lowered chandelier, the other person will raise it so you can continue along the parkour path.

You will get to another chandelier, this time your partner will have to lower it so you can hop on then raise it so you can continue along the path.

You will get to another chandelier that will need to be lowered, then raised. This will be the last partner action since the skull is on the chandelier after it.

Solo parkour

Get to this area, and go in the building on the right of the second picture.

Once inside, turn the valve and exit. We will then move into the nook with a bunch of trash

Once we are in the trash corner, we will climb the dumpster and barrels to get to the walkway

Follow the walkway, turn left, and pick up the power cell.

Put it into the glowing receptacle, and hit the lever beside it.

Once the lever has been switched, the door inside the building should be powered on. Go inside, press the button and just past the door will be the skull

As an additional note per ΛQ
"There is a cheese specifically for Ogryn: You can hop onto the barrel just under/near the power cell and jump up to grab it easily through the hole in the platform it's nestled in without climbing the whole thing."

Solo puzzle

Proceed through the map until just before the bar just before the midevent. There will be an alley outside of it, with a dead body and a key.

After getting the key, go inside the bar and go to the left side. There will be a locked gate that you can now open with that key.

Go through the hallways until you get to a breachable wall, breach it and grab the key

Exit this section, and return to the bar. Behind the bar is another chainlink gate. Open it and the skull is at the back.

Solo Parkour

Credit: The AndersBot for pointing it out to me

This skull is after the second medicae in the area with the long staircase that leads to the cellar.

On the right upon entering the room with the staircase there will be a broken railing, this is where we start the parkour.

It is a quick and simple parkour, jump towards each of the indicated platforms. You could sprint jump to make it easier, but it should be unnecessary. Hold space after jumping to automatically mantle just in case you fall a little short.

Towards the last jump, there is a pipe on the left in the next picture, above the teammate ui, that you could jump on as well to make it easier (credit: =M.D= Lion). The skull will be next to the brightly glowing candle